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Plants & People: The Intersection of Botany and Human Experience

Aronia melanocarpa: fine wines to phenols

Aronia melanocarpa, also known as Aronia Berries or Black Chokeberries, recently garnered attention for its potential as a “superfood” that combats a variety of diseases. Its scientific name refers to its dark pome fruit, while its common name is a reference to its tart and astringent flavor. Originally native to woodlands and bogs in North America, this shrub has historically been used as medicines and food by the Potawatomi and Abnaki people. It was first described as “Mespilus arbutifolia var. Melanocarpa” in 1803, by the French botanist André Michaux. It was then recategorized as “Aronia melanocarpa”, by American botanist Stephen Elliot in 1821 .Now, it is enjoyed throughout Eastern Europe in jams and wines after its introduction post World War II. Beyond commercial production, this plant can be found planted in rows in people’s gardens. These plants are deciduous, with green, gold leaves that change color to a burgundy red in the fall. They produce inflorescences of small white flowers that ripen into the berries that provide many health benefits. Recent research into the uses of bioactive compounds such as antioxidants, anthocyanins, proanthocyanidins, flavonols, and phenolic acids suggests that extracts from the plant could be useful for anti-inflammatory purposes or treatments for diseases ranging from diabetes to cancers. As a result, the industries for Aronia berries have been on the rise and these berries have been introduced into many new recipes for everyday use. By Natalie Keung and Joanne Kwak #chokeberries

Aronia melanocarpa, commonly known as the Aronia Berries or the Black Chokeberry, is a deciduous shrub cultivated throughout Eastern Europe. Originally indegenous to North America, the plant was known as nîki’mînûn or sakwako’mînûn by the North American Potawatomi and Abnaki tribe. When European settlers arrived in the Americas they named the plant Mespilus arbutifolia var. Melanocarpa in 1803, by the French botanist André Michaux. It was then recategorized as “Aronia melanocarpa”, by American botanist Stephen Elliot in 1821 . The genus name, Aria, is derived from the Greek word “aria”, while the species name, melanocarpa, is derived from the words "Melano"- meaning black, and "carpa", meaning fruit. Commonly known as the Chokeberry, this name is a reference to the berry’s tart flavor. As one ethnobotanical book from 1933 noted, “the natives eat the berries from this plant but they are entirely too bitter to suit the white man”. However, the plant is now used in a variety of foods throughout Europe (Englels et.al 2014) .

The Aronia berry is a member of the Rosacae family, or the “Rose” family, which includes plants such as roses, apricots, or almonds. The plant is one of three plants within the genus Aronia, and is closely related to Aronia arbutifolia, the red chokeberry, and Aronia prunifolia, the purple chokeberry, which some consider a hybrid of the two species instead of a fully fledged species itself. Although originally these plants were thought to be part of the Photinia genus, a 2004 study proved that genus Aronia was instead a distinct lineage and not closely related (Englels et.al 2014) . In modern cultivation, individual specimens of Aronia melanocarpa used in cultivation are often very closely related to each other, as new plants are established from cuttings although there are new efforts to produce new lineages that will have higher nutritional content.

Originally, this plant was most commonly found in the Northeastern part of North American, running from the Great Lakes region to the Appalachian Mountains. It was most commonly found in swamps, wet thickets, or in the margins of ponds and lakes, although it could also survive in drier soils. The plant is known to be hardy and does not suffer from any major pests or disease. The plant acts as a good source for birds and their flowers produce pollen to insect pollinators, with their main pollinators being small bees. On average the plant lives upwards of 25 years, depending on how favorable the location is. Due to its hardiness, it is known to be invasive in Eastern Europe (Henrich, 1976) .

Traditionally, the berries were used by North American Forest Potawatomi and Abnaki tribes in preparation of pemmican, a high calorie food made from animal fat, dried powdered meat, and sometimes fruit. Also, they used the fruits for medicinal teas, make dyes, cure meats, and as nutritional dried treats in the long winter months. The berries and the bark were used as an astringent (Engels et.al 2014).

After World War II, the first shrubs of Aronia melanocarpa were brought to Europe and Russia, specifically Altai, Moscow, and St. Petersburg. Starting 1946, large- scale black chokeberry cultivation was introduced to the former Soviet Union, including Belarus, Moldova, Russia, and Ukraine. In 1976, aronia was introduced to Japan and by the 1980s, it was introduced for cultivation in Bulgaria, Czechoslovakia, East Germany, Poland, Slovenia, Denmark and Finland. Poland was the main supplier of chokeberry to Germany despite cultivation being present in East Germany since 1976. In 1996, Jan Mills, president and CEO of Artemis International Inc. took the main commercial cultivars from Polish agricultural schools and brought them back to Indiana. Since then, the chokeberry bush is used as an ornamental plant in the United States. In England, it received the Royal Horticultural Society’s Award of Merit in 1972 for their white flowers in the spring, green foliage in the summer, and fruits in the late summer and autumn (Engels et.al 2014).

Aronia melanocarpa is a popular shrub on the industrial and amateur scale in Central Europe (Engels et.al 2014). It is used as an ornamental in the US and the berries are eaten fresh or incorporated into cooking in various locations and industries. In Poland, black chokeberries are used on a large scale by the food, pharmaceutical, and cosmetics industries. In Lithuania, these berries are used to produce wine. In general, Aronia is used in juice, jams, teas, and cordials. In the canning industry, Aronia are used to produce compotes, confitures, fruit jellies, marmalades, and the juices from the fruit are used to adjust or enhance flavor along with coloring. The fresh pressed juice is used as raw material for the manufacture of alcoholic sparkling wines, dessert wines, and liqueurs. In dairy processing, ice cream, yogurts, cream desserts, and sorbets include the berries. Black chokeberries are used for baking, specifically for praline fillings, cake fillings, fruit bars, cereals, and gummy bears. The pomace, or solid remains of the fruit after pressing, are used as color pigment (Engels et.al 2014).

In Eastern Europe, thousands of Aronia melanocarpa are grown commercially. In addition to commercial production, Aronia melanocarpa is also sold as an ornamental. In the U.S., the aronia berry industry is still developing, but currently is focused more in Harrison County inWestern Iowa. ‘Autumn Magic’ and ‘Iraqis Beauty’ are the two most commonly sold ornamental cultivars of Aronia melanocarpa. Different cultivars have different purposes. ‘Viking’ is a vigorous, productive variety from Scandinavia, which can grow to a height of 6 feet. ‘Nero’ is a shorter growing variety, reaching a height of 3 to 4 feet, with dark blue berries. In the U.S., a selection from a native source in Michigan is being sold as ‘Morton’ black chokeberry. It is marketed in the Midwest under the trademark Iroquois Beauty™. ‘McKenzie’ is a larger form that grows from 6 to 12 feet developed especially for conservation use and agroforestry benefits by the Plant Materials Center at Bismarck, North Dakota (Ochmian 2012).

The Aronia melanocarpa shrub produces their first fruits in 3 years and the first large harvest occurs after 5 years (Mahoney 2019). The berries tend to hang in clusters of up to 12 berries. They are harvested by hand or mechanically with a blueberry picker in late August or late September after they ripen (University of Maine 2022). This shrub has many adaptations from their fibrous roots. Aronia berries can grow without irrigation because these fibrous roots store moisture. These roots also prevent soil erosion, which protects the soil. This shrub grows and spreads by suckering, allowing it to reproduce easily. However, when growing commercially or in personal gardens, cultivators tend to plant the Aronia in rows in slightly acidic soils for the best results (University of Maine 2022).

Aronia melanocarpa are deciduous, rounded, but upright shrubs that become leggy as they age. These shrubs grow to a height of 2-3 meter but vary based on the particular cultivars (Englels et.al 2014). The bark of this shrub is thin and smooth. The leaves are simple and alternately arranged with serrated edges and an elliptical shape. Since the Aronia melanocarpa are deciduous, the leaves change color in the fall. Throughout the year, the leaves are gold or yellow and dark green but as fall approaches, the leaves turn into a burgundy or red color (N.C. State University 2022). New branches of this plant range from grey to brown with scattered lenticels and are often covered in small white hairs, which smooth and become covered with a white film with age. New buds are sharp-pointed with a prominent terminal bud and smaller lateral buds pressed against the twig.

In late May to June, the plant produces 20-30 small white flowers that eventually mature into dark purple berries that span in diameter from 6 to 13 millimeters and weigh from .5 to 2 grams, similar to the size of blueberries. The small white flowers are perfect, with both male and female reproductive structures. The inflorescences and each have 4-5 white petals with pink anthers. They are protogynous and self compatible and reproduce apomictically via gametophytic apomixis, which results in embryos identical or nearly identical to maternal plants (Mahoney 2019). The primary pollinators of these shrubs are small bees. Reproduction in the wild is mainly by seeds, which are small, slightly more than 1/16 of an inch. These seeds must be cleaned and dried after harvest for the best germination. However, in cultivation the plant is propagated by cuttings (Mahoney 2019).

Historically, the Black Chokeberry was used both as food and medicine by a wide variety of people. Especially in Eastern Europe, the food is a common flavoring and colorant. Currently, in addition to its use as an ornamental shrub and food source, this plant has drawn attention for its high levels of antioxidants, anthocyanins, proanthocyanidins, flavonols, and phenolic acids (Halina et.al, 1970). A variety of studies are being conducted on the impact on these compounds on reducing the risk of non-communicable diseases such as diabetes, metabolic syndrome, and neurological disease, acting as an anti-inflammatory, or preventing cancers (King et. al, 2020) (Martin et.al, 2014). Current research is being conducted into how to best breed new strains to maximize the concentration of bioactive compounds, how to process the berries to retain bioactive compounds as well as how to best apply these compounds to treatments in a clinical setting (Tolic et.al, 2015). These promises have categorized the plant as a “superfood”, and a variety of supplements are on the market, although not approved by the FDA for medicinal use. Anecdotal evidence claims that these supplements have helped reduce inflammation, reduce blood sugar levels, and reduce blood pressure.

Notes

Because this plant is not very commonly consumed, at least in the Americas, it was quite difficult to find reputable english-languages sources. Most large databases lacked images or records on the Aronia genus as a whole, much less the specific species melanocarpa. Many sources found online differed in their claims. For example, one source claimed that the plant was common in East Asia after its introduction after the Soviet Era, while a Smithsonian painting by a Japanese master dated a depiction of the genus to between 1797-1858, or the Edo period. Other sources claimed that the plant was not prevalent in Eastern Asia at all. Additionally, many sources very clearly had plagiarized from each other, and neglected to cite sources. Overall, exploring this plant helped us think more critically about the information we found online, and helped us appreciate the effort put into clear and reputable sources.

References
A. PERSSON HOVMALM, H. E. L. E. N. A., Jeppsson, N., V. BARTISH, I. G. O. R., & Nybom, H. (2004). RAPD analysis of diploid and tetraploid populations of Aronia points to different reproductive strategies within the genus. Hereditas, 141(3), 301-312.
“Aronia Melanocarpa.” Aronia Melanocarpa (Black Berried Aronia, Black Chokeberry) | North Carolina Extension Gardener Plant Toolbox, https://plants.ces.ncsu.edu/plants/aronia-melanocarpa/.
Culture of aronia for fruit production - cooperative extension: Agriculture - University of Maine Cooperative Extension. Cooperative Extension: Agriculture. (n.d.). Retrieved May 10, 2022, from https://extension.umaine.edu/agriculture/aronia/culture/
Engels, Gayle, and Josef Brinckmann. “Black Chokeberry Aronia Melanocarpa Family: Rosaceae.” Black Chokeberry - American Botanical Council, https://www.herbalgram.org/resources/herbalgram/issues/101/table-of-contents/hg101-herbpro-chokeberry/.
Everhart, E. (n.d.). Aronia - a new crop for Iowa. News. Retrieved May 10, 2022, from https://www.extension.iastate.edu/news/2009/mar/110401.htm
Heinrich, B. 1976. Flowering phenologies: Bog, woodland, and disturbed habitats. Ecology. 57(5):890-899.
Halina Maria Ekiert, Agnieszka Szopa, & Paweł Kubica. (1970, January 1). High production of depsides and other phenolic acids in different types of shoot cultures of three Aronias: Aronia melanocarpa, Aronia arbutifolia, Aronia × prunifolia. SpringerLink. Retrieved May 9, 2022, from https://link.springer.com/referenceworkentry/10.1007/978-3-030-30185-9_11
King, E. S., & Bolling, B. W. (2020). Composition, polyphenol bioavailability, and health benefits of aronia berry: A review. Journal of Food Bioactives, 11.
Mahoney, Jonathan D., et al. “Sexual and Apomictic Seed Reproduction in Aronia Species with Different Ploidy Levels.” Hortsci, American Society for Horticultural Science, 1 Apr. 2019, https://journals.ashs.org/hortsci/view/journals/hortsci/54/4/article-p642.xml.
Martin, D. A., Taheri, R., Brand, M. H., Draghi II, A., Sylvester, F. A., & Bolling, B. W. (2014). Anti-inflammatory activity of aronia berry extracts in murine splenocytes. Journal of Functional Foods, 8, 68-75.
Tolić, M. T., Jurčević, I. L., Krbavčić, I. P., Marković, K., & Vahčić, N. (2015). Phenolic Content, Antioxidant Capacity and Quality of Chokeberry (Aronia melanocarpa) Products. Food technology and biotechnology, 53(2), 171–179. https://doi.org/10.17113/ftb.53.02.15.3833
Ochmian, Ireneusz, et al. Notulae Botanicae Horti Agrobotanici Cluj-Napoca, Comparison of Some Morphological Features, Quality and Chemical Content of Four Cultivars of Chokeberry Fruits (Aronia Melanocarpa).
“WSU Clark County Extension.” Black Chokeberry - Aronia Melanocarpa - PNW Plants, http://www.pnwplants.wsu.edu/PlantDisplay.aspx?PlantID=59.

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1. Lepidium meyenii: The fertility root Maca… maybe you’ve heard of it, maybe you consume it on a daily basis. Maca is the highest elevation crop known today, and is native to the high elevation Andes mountains in Peru, growing in elevations up to 4500 m high! Maca has been a highly valued commodity for many indigenous peoples within the Andes mountains of Peru throughout history and persists to be valued today. Known for its turnip-like shape due to its bulbous hypocotyl, it is highly sought after for its roots. Consuming the roots will result in many notable benefits, such as nutritional benefits, mood improvements, and enhancements in energy, strength… even fertility, all which are backed by numerous studies. These benefits are all due to the numerous nutrients, fibers, bioactive compounds, and natural compounds lepidiline A and B that are contained within Maca. Combined, they all make Maca the nutritional powerhouse that is today. Once this nutritional powerhouse was advertised to the world, it became an immediate hit. The demand for Maca created a market boom and countries started trying to cultivate the plant to meet the demand. One country that cultivated Maca is China. In the high mountains of Yunnan, Maca was successfully cultivated and thus became a common ingredient in Chinese traditional medicine. As such, Maca remains an important dietary source not only in Peru, but as well as worldwide! With more research being done today to further explore what components are in Maca, only time will tell what additional health benefits Maca has to offer us. By Mellanie Gamero and Jacqueline Duong
2. Moringa oleifera: Domestication Histories and Emerging Deviations from Indian Origin The superfood, Moringa oleifera, has a rich history deeply rooted in its native Indian domestication and usage. Also widely known as “The Miracle Tree”, M. oleifera is often recognized by its white flowers and long-seed pods that are greatly sought after. Not only does M. oleifera provide an incomparable food source and multipurpose oil through its pods, but all parts of the plant may be used and consumed. The versatility of this plant is venerated in Indian culture where its consumption and, most importantly, medicinal usage involve all parts of the plant. Due to its spread across other countries, the species and its utilization have evolved to suit individual cultures and their own domestication desires. In the Philippines and North and South Africa, its nutritional value is greatly relied upon to treat malnourishment, purify water, and support community infrastructure while the Indian native medicinal usage is less emphasized and not well known. The Americas, on the other hand, have increasingly toyed with the novel M. oleifera ‘organic-superfood’ craze that tends to remove emphasis from its cultural value and multipurpose utilization to instead fawn over its supposed energy boosts, skin care implementations, and uses in its powder – likely less effective and increasingly more expensive than the original. Though its original purpose grows increasingly colonized, the domestication ventures globally of this superfood continue the legacy of the Moringa oleifera species itself – marked genetic increases in energy production associated with plastic to nucleus gene duplications. As each nation tends to and selects for various aspects of this energy contribution – whether for the nutritional, medicinal, or vanity value – the variation in these M. oleifera may grow towards eventual speciation. By George Vetushko and Lauren Guevara #drumsticktree
3. Salvia hispanica: A Source of Strength Traditionally a key component in Aztec and Mexican culture, chia seeds have been used for centuries for their nutritional, decorative, and medicinal qualities. A common staple in many cultures around the world, these tiny seeds pack a powerful punch and are enjoyed by fitness enthusiasts everywhere. While the seeds themselves are easily recognizable, less well known is their source: a plant known as Salvia hispanica. This flowering, annual herb has a fascinating history and distinctive morphological traits that have evolved over time due to human cultivation. By exploring the myriad ways that this plant has interacted with people over the course of history, we can gain a greater appreciation of how chia seeds have been an important source of strength and inspiration for many. By Joshua Matsuda, Sarah Hoffman, Julia Lindner #Chia
4. Breadfruit: The Intrepid Fruit's Journey with Indigenous Peoples and Colonization Breadfruit (Artocarpus altilis (Parkinson) Fosberg) is an intrepid dicot with a rich history and culture surrounding it. Breadfruit trees are a single-trunked tree with a spreading, evergreen canopy. Breadfruit trees are monoecious with both the male and female flowers on the same tree. The male flowers are club shaped, and the female flowers are thousands of small flowers with 2 anthers around a spongy core. The name is derived from the Greek words bread and fruit (artos = bread & karpos = fruit). There are over 2,000 documented names because of its widely spread distribution across the Pacific. A. altilis originated from the South Pacific and was later spread throughout Melanesia, Micronesia, and Polynesia by voyagers. As a result, breadfruit is adapted to a wide variety of climates and conditions in equatorial lowlands. In addition, breadfruit trees thrive in humid tropics and can grow in a variety of soils. Breadfruit became a staple crop because of its nutritional and medicinal value. In the 1700s, breadfruit’s ease of maintenance and nutritional value made it a prime crop, in the eyes of colonizers, to feed African slaves in the Caribbean. Today, breadfruit continues to have an important role in sustainability, fighting malnutrition, and increasing food security. There are research efforts to increase the nutritional value and initiatives to promote breadfruit cultivation in tropical and subtropical regions. For example, The National Tropical Botanical Garden established the Breadfruit Institute in 2003 in hopes of promoting the study of breadfruit for agroforestry, reforestation, global food security and economic development. #Angelica Soriano and Bucky Squier
5. Tarwi: A forgotten gift from the Andes Tarwi is an Andean lupine that has been cultivated by those native to the Andes for at least 1500 years. The scientific name of this lupine is Lupinus mutabilis, Lupinus meaning “wolfish,” and mutabilis meaning “variable.” It can survive in high elevations ranging from 1650 to 3300 meters. This plant is a dependable crop that has been used by the pre-Inca and Inca people in crop cycling with other native Andean plants such as quinoa and potatoes. Documentation of the use of tarwi has been recorded archeologically in Naza culture tombs and Tiahuanaco ceramics. There are also records of high cultivation of tarwi through Spanish taxation records that recorded 10,000 hectares cultivated. The use of tarwi decreased following Spanish colonization when Old World crops were introduced to the area; by 1980, only 5,200 hectares of tarwi were cultivated in Peru. Tarwi has colorful purple flowers and palmate leaves, making the plant strikingly beautiful. The tarwi plant can self-pollinate and does not rely on one specific pollinator making it less vulnerable to fluctuations in pollinator populations. In addition to being dependable in pollination, tarwi can be planted in poor soil because it can fix nitrogen in the soil. The tarwi plant contains alkaloids, making it less susceptible to pathogens and pests. The seeds of this plant are consumed, but the seeds need to be soaked in water for two to four days to remove the water-soluble alkaloids that make the seeds bitter. As an Andean native plant, Lupinus mutabilis has various properties and uses that have enhanced the diets and lives of Peruvians and other South Americans. By Lysol Patino and Sarah Simpson #tarwi
6. Smallanthus sonchifolius: The Superfood of the Andes Smallanthus sonchifolius is a plant that in recent years has gained a lot of traction internationally. Its ability to store sugars in the form of carbohydrates has resulted in a very profitable market for yacón cultivators. People have begun looking at this food as a healthier alternative to sugars which has in turn led to its global expansion. A food that was once limited to exist within the Andes has now expanded to other continents. Yacón's human impact has been a part of South American culture for thousands of years, as it is believed to have originated before Incan times. During this time period yacón's consumption was limited to religious events; today however, it is a food that is consumed globally. Apart from its sweet tuberous roots, other parts of the plant that are edible are the leaves and the flowers. Together these attributes allow yacón to be known as one of the superfoods from the Andes. By Jessica Manriquez and Amanda Lin #yacón
7. Irvingia Gabonensis: The Untapped Diversity of Dika Plant Irvingia gabonensis is a multipurpose, undomesticated fruit tree native to West and Central Africa. Belonging to the family of Irvingiaceae, this evergreen tree species can grow to a height of 40 meters and attain a diameter of about 120 centimeters. The tree produces mango-like fruits that are well known for their nutritious kernels. Even though the fruit of this plant is also referred to as the African bush mango, the plant has no connection to the actual mango fruit. The dika tree can be found in the dry and wet tropical zones such as Nigeria, Angola, Uganda, Congo, Cameroon, Ghana, Togo, and Benin. It is also known to grow in farmland areas, semi-deciduous forests, and gallery forests. Despite its obscurity on the international level, this species should hardly be considered a minor resource. Almost all parts of the plant have a use from its fruit to its bark. Additionally, it serves as a main source of food and income for indigenous groups in Africa such as the Baka tribe from Central Africa. Due to how prized this plant species is, the dika plant is left untouched by the indigenous people when forests are cleared. Once domesticated, the plant can potentially be used to treat the high levels of malnutrition in Africa. Yet, with increasing local and regional demand for the plant’s kernels, the African bush mango has been facing intense exploitation. Additionally, the disappearance of local wildlife such as elephants are affecting the plant’s dispersal. As a result, the I. gabonensis plant has been classified as a highly endangered species. By Ian Morris and Nikhitha Nair #dika
8. Goji Berries: The Key to Eternal Youth? Goji Berries have become a symbol of the Superfood movement which has continued to gain traction in the US and across the world. Although they have served as a token of this emerging trend, goji berries have been renowned in Asia for their nutritious qualities for many centuries. Its use has been documented for almost 2000 years in traditional Chinese medicine (Gross et al., 2006). The goji industry has skyrocketed during the 21st century as they’ve been recognized for their nutritious qualities. Goji berries have a rich cultural history which has contributed to their claim to fame. In 2003, health writer Earl Mindell distributed a booklet which suggested that goji berries had anti-cancer properties - this drove their entry into the Superfood market. His claims were based on the myth of Li Qing Yuen; the herbalist supposedly consumed goji berries daily which allowed him to live 256 years (Earl Mindell and Rick Handel, 2003). While Mindell’s booklet was full of false exaggerations, there is merit to many of his claims regarding the nutritional benefits of goji berries. Two species are colloquially known as goji berries: Lycium barbarum and Lycium chinense. Their unique vegetative and reproductive morphology makes harvesting the berries labor intensive. Both the goji berry and immature leaves from L. chinense and L. barbarum are eaten by many for their health benefits - while the benefits have been known for centuries, the underlying macromolecular structure that dictates these properties is only being explored in more recent decades. By Catherine Nordstrom & Ismael Rodriguez #Goji
9. Tiger Nuts: Cultivar, Weed, and the Plasticity of Plant-People Relationships Contrary to its name, the tiger nut is the edible tuber of Cyperus esculentus, a sedge that originates from the Mediterranean area. While it was considered a valuable crop in pre-dynastic Egypt where it was even entombed with Egyptian officials, Cyperus esculentus became widely known as an invasive weed during the 19th century. Because of its low yield, it was never considered valuable enough to cultivate as a cash crop. Once again, however, the tiger nut is regaining its popularity but now as a superfood. With its high fiber content and ability to promote gut health, it continues to rise in popularity as one of many superfoods available in the market today. By Phan Tran, Karla Gonzalez
11. Aronia melanocarpa: fine wines to phenols Aronia melanocarpa, also known as Aronia Berries or Black Chokeberries, recently garnered attention for its potential as a “superfood” that combats a variety of diseases. Its scientific name refers to its dark pome fruit, while its common name is a reference to its tart and astringent flavor. Originally native to woodlands and bogs in North America, this shrub has historically been used as medicines and food by the Potawatomi and Abnaki people. It was first described as “Mespilus arbutifolia var. Melanocarpa” in 1803, by the French botanist André Michaux. It was then recategorized as “Aronia melanocarpa”, by American botanist Stephen Elliot in 1821 .Now, it is enjoyed throughout Eastern Europe in jams and wines after its introduction post World War II. Beyond commercial production, this plant can be found planted in rows in people’s gardens. These plants are deciduous, with green, gold leaves that change color to a burgundy red in the fall. They produce inflorescences of small white flowers that ripen into the berries that provide many health benefits. Recent research into the uses of bioactive compounds such as antioxidants, anthocyanins, proanthocyanidins, flavonols, and phenolic acids suggests that extracts from the plant could be useful for anti-inflammatory purposes or treatments for diseases ranging from diabetes to cancers. As a result, the industries for Aronia berries have been on the rise and these berries have been introduced into many new recipes for everyday use. By Natalie Keung and Joanne Kwak #chokeberries
12. Eragostis tef: From Injera to fertility and everything else in between Teff is a unique member of the grass (Poaceae) family that is a staple in East Africa and accounts for a large portion of East African diets. Teff, known for being the plant with the smallest seeds, is a labor intensive crop that is necessary in Ethiopian and Eritrean cuisines. This necessity is born from the traditional usage of Teff flour being used in the fermentation and creation of Injera, a sour spongy flatbread. This demand of Injera, and therefore Teff flour, has led to 31% of Ethiopian landmass being used for the cultivation of Teff. The agronomics of Teff are interesting in that Teff can only thrive in climates similar to the East African Highlands and the planting and harvesting of Teff are known to be labor intensive due to its rooting pattern and its small grain size. Teff itself has multiple physiological variations ranging from varying colors for both the seeds and the plant to variations in Amino Acid, Protein, and nutrition profiles. Teff’s stability as a choice crop in East Africa is accredited to its hardiness to survive drought and water logged conditions when rooted in the proper soil conditions, shelf stability, and the straw can be used for livestock forage feed. Teff is thought to originate some time around the 6000 B.C, but cultivated at some point around 2500 B.C as the earliest known record of Teff, is seeds found in bricks of the Dahshur Pyramids, most prominently the Red Pyramid of Sneferu. By Luis Angel Yepez #tef
13. Sacha Inchi: Super Seeds Plukenetia volubilis, commonly known as sacha inchi, is extremely valuable as both a source of nutritional and medicinal benefits. Sacha inchi has been cultivated for thousands of years in the Amazon Basin by the Incas and other indigenous peoples, who have used it as both food and medicine. The incredible properties of this plant are primarily derived from the seeds and the oil extracted from them. The seeds contain very high levels of essential polyunsaturated fatty acids, including both omega-3 and omega-6, which have been shown to promote cardiovascular health. The seeds additionally have high protein content and have been used in traditional Peruvian cooking as a nutritious staple. A number of dishes are traditionally prepared with sacha inchi, including snacks, confections, soups, and porridges. Compounds found in the seeds have been used by indigenous groups to treat a variety of illnesses, including rheumatism, muscle pain, high cholesterol, cardiovascular issues, and gastrointestinal ailments. Scientific studies of compounds derived from sacha inchi have suggested that the plant may be useful in suppressing tumor proliferation and treating dermatitis, chronic inflammatory diseases, and cardiovascular disease. The plant has also been found to have antioxidant properties, which may promote health and support immune function. The versatility of the seeds and seed oil derived from sacha inchi as both highly nutritious food and extremely powerful medicine encourages further study and cultivation of this fascinating plant, which is not fully domesticated. The high genetic diversity of wild sacha inchi plants offers a wonderful starting point to cultivation efforts, with the potential for increasing yield and optimizing nutrient levels. By Amanda Bueno-Kling and Giselle Correa #sachainchi
15. Açaí Berry: The Exploitation of Labor in South America The açaí berry is a small drupe, similar in size to a blueberry but deeper purple in color. Native to Eastern Amazonia, the açaí berry has only recently begun to be popularized as a “superfood” in urban communities in the United States within the past 20 years. This superfood claims to be helpful for a variety of major health concerns including, but not limited to, arthritis, weight loss, high cholesterol, and detoxification (NCCIH). Even though research is limited and these claims remain unproven, the popularity of açaí berries still manages to grow exponentially each year. The global market for the açaí berry is expected to exceed $2 billion by 2026 (McCoy). The only proven claim than can be made about the açaí berry is that its rise in popularity only serves to threaten the lives of families, both adults and children, that are living in the surrounding communities in the Amazon who are responsible for harvesting the berries and are, in turn, routinely exploited for cheap, quick labor. More often than not, these açaí pickers, known as peconheiros, are forced to work in critically dangerous conditions and are paid little to nothing. Peconheiros are regularly exposed to serious injuries including bone fractures, accidental knife wounds, and venomous snake and spider bites. With the açaí trees growing upwards of 50 feet tall and only a braided rope being provided to climb, falls are common– and sometimes fatal. Further, peconheiros have no way of advocating for themselves in the event that they do become injured in the forest. No laws or regulations exist that protect the safety and wellness of the peconheiros. by Roberto Vindel, Pauline Le, and Max Kwon #Açaí
16. Carob: From Pod to Pantry Ceratonia siliqua, commonly known as carob, St. John’s Bread, and locust bean, is a flowering deciduous tree in the legume family that produces large seed pods with a wide range of commercial uses. Carob has existed long before the start of agriculture, with origins across the Mediterranean basin dating back to the Paleolithic age. Recent phylogenetic data suggests domestication of carob occurred through localized selection of wild genotypes, with some long-distance westward dispersal events by the Romans or Arabs. Carob trees are valued ecologically for their low maintenance, high adaptability to poor soil conditions, and drought resistance, with some even living for 100 years or more. They feature thick, dark green and broad foliage, thus ornamental trees can be found throughout California, Australia, and other parts of the world with Mediterranean climate. However the main attraction of Ceratonia siliqua is its brown, leathery pod. The carob pod can be split into two main components: the pulp and the seeds. The naturally sweet pulp has a sugar composition of up to 48-56%, though it also contains cyclitols, fiber, and polyphenols such as tannins. Carob pulp is a nutritionally significant source of amino acids, minerals, and vitamins. The seeds contain a galactomannan gum also known as carob bean gum (CBG) or locust bean gum (LBG) used as a thickening agent in many food products. Carob pods are associated with the prevention and treatment of a wide variety of diseases, including diabetes, hyperlipidemia, irritable bowel syndrome, and colon cancer. In the past few decades carob has been marketed simply as a healthier chocolate alternative, but its rich history, health benefits, and potential applications in areas affected by climate change should allow it to stand on its own as a staple in the pantry and beyond. By: Chloe Fuson and Skylar Yee #Carob
17. The Wonder Berry : Phyllanthus emblica Known for being rich in both nutrition and history, the amla berry or as Carl Linneaus named it in 1753, Phyllanthus emblica, can be classified as India’s “Wonder Berry.” Originally believed to come from the semi-arid areas and plains of Northern India, the plant is native to Asia's tropic south-eastern regions such as Vietnam, Laos, and Thailand. Such regions provide conditions like mixed forests and elevations, that the plant cannot live without. However, with the fruit’s beneficial properties being so popular, countries all over the world cultivate the amla berry with India well beyond 100,000 acres of amla development. The amla berry’s influence has even reached as far as the United States where seed distribution first began in the early 1900s. It is said that the Indian god Brahma’s tears germinated the first seeds. In fact, the origin of the fruit’s usage can be traced hundreds of years ago where it was and still used today as a healing remedy. This fruit has been used to cure fevers, joint inflammations, and constipation for hundreds of years. Even recent research has suggested that the antiviral and antibacterial properties of the amla berry may be able to prevent cancer, diabetes, and excessive cholesterol. Despite only being the size of a golf ball, the amla berry is very high in vitamin C, vitamin E, vitamin A, iron, and calcium. In contrast to the size, these berries grow on trees that can reach heights of 12 meters all while surrounded by the beauty of greenish-yellow and pink flowers running across the branchlets. Ultimately, it is safe to say that such a plant is truly a wonder. By Natalie Nartz and Matthew Nguyen #amla
18. Annona cherimola: The most delicious fruit known to man Cherimoya is a fruit from the genus Annona, it is cone shaped, green, leathery skin with a creamy and sweet flesh. The fruit has a sweet taste and it is rich in nutrients and minerals. Due to its creamy textures, it is also called the custard apple. It is believed to be native to the Andes Mountain range in South America but it has been distributed and planted in different areas of the world today. One of the major problems for cherimoya cultivation is inadequate natural pollination since the male and the female structures of each flower do not mature at the same time. Due to its rich mineral content cherimoya is known to have some health benefits including lowering blood pressure. However, cherimoya and other Annona species contain annonacin which is a toxin that can affect the nervous system and the brain. Some studies have suggested that high amounts of consumption of Annona fruits can lead to increased risk of a specific type of Parkinson’s disease that does not respond well to medication. The seeds can be toxic, so when eating the fruit, it is best for the skin and seeds to be discarded. While Cherimoya has lots of health benefits and can be delicious it needs to be eaten in moderation to avoid its negative effects on the nervous system. by Shiva Nia, Kyi Ther Min #Cherimoya
19. M. Citrifolia: A Distinctive Fruit with Exceptional Traits M. citrifolia is a unique, flowering and fruit bearing plant that goes by many names such as the most notable, Tahitian Noni fruit, Indian Mulberry or Great Morinda. It has many deeply rooted cultural histories in many regions of the world because of its experiences in many trade avenues and nutritional and medicinal applications. The plant is most easily identifiable, as some of its names suggest, by the distinctive fruits it grows, a green or white, depending on the ripeness, typically mango sized, that is said to have a strong odor similar to cheese and also has a bitter taste. Although it may not be the most attractive plant in many senses, it has been found to be very useful with almost all parts of the plants, including the roots, leaves, seeds, bark, shoots, being used in various ways, whether it be for wellness, nourishment or trade value. Nowadays it is most often found in regions of Oceania, the areas of Polynesia, Micronesia, and Melanesia, due to the favorable conditions of the tropical environments, as well as the agricultural practices of the Pasifika peoples over the past few thousand years. As a consequence of the now main homeland of M. citrifolia being the islands of the Pacific Ocean, discussions of the origination of the plant can sometimes be intricate, although some discourse and evidence can point to a certain continent as the place of emergence. Overall M. citrifolia has been utilized by numerous peoples and communities and continues to be a key element in several of these life ways. By: Megan Chen and Kokonow Kinney #noni
20. Pouteria lucuma: An exploration of historical and modern medical uses of the Andean lucuma fruit Pouteria lucuma is a fruit tree of the family Sapotaceae of the order Ericales native to the Andean Valley of South America. The fruit is commonly used in Peruvian desserts, especially icecream, for its unique flavor, often described as resembling maple, butterscotch, or sweet potato. It was used as a subsistence crop by the native peoples of coastal Chile and Peru as early as 700 BC. The fruit was an important part of fertility rituals of these peoples, as its consumption was thought to promote lactation in new mothers. No studies have been done to that effect, however there has been significant research into other medicinal properties of the fruit. The oil extracted from the seeds and skins of the fruit, which are typically discarded as industrial waste, has high phenolic content and has been shown to aid in wound closure and skin regeneration. These properties have made the fruit of special interest to the skin care industry. By Jasmine White, Quinn Schwabauer, Jin Zhu #lucuma
21. Avocado: New World Nutient Pack The avocado has extensive cultural and anthropogenic ties to human society, with three main cultivars representing the three different domestication and cultivation practices for the crop. The Mexican, Guatemalan and West Indian races (P. americana var. drymifolia, P. americana var. guatemalensis, and P. americana var. americana) are ancestors to more than 30 types of modern avocado cultivars, and have linguistic and historical ties to Mesoamerican cultures that domesticated them at different geographical locations and time periods. Not surprisingly, the different domestication and cultivation practices of the three ancestor cultivars have led to different optimal abiotic conditions for different types of avocados, with Latin America and Africa being the main suppliers for the majority of cultivars that enter the global market. Moreover, all parts of the plants can be used, although the most common part of the plant consumed is the fruit. The leaves and seeds are sometimes used as medicine, and the oil extracted from the fruit can also be used in medicine and cooking. Additionally, the avocado’s high concentration in fiber, folate, antioxidants, potassium, vitamin E, and magnesium per ounce makes this crop a superfood, and is most commonly used as an ingredient for guacamole. Modern society recognizes the importance of avocados, especially considering that we are entering an age where high-caloric foods are readily accessible and easier to purchase than healthy foods. As such, cultivation practices are also shifting from large-scale industrial productions that use heavy pesticides to smallholder farmers who can implement traditional cultivation practices that are more sustainable for the farmers and the ecosystems around these farms. By Amanda Leyel and Hojae Lee #Avocado
22. Camu Camu: The Super Fruit of the Amazon Basin Myrciaria dubia (Kunth) McVaugh or camu camu, is a tropical shrub-like tree that is best-known by its highly acidic, nutrient-rich berries. The plant has been extremely important for the Yanomamo and Kayapo Indigenous peoples of the Amazon Basin, who have harvested them for thousands of years. Wild and domesticated populations alike are harvested by guiding canoes through the branches during the wet season. The camu camu plant grows to be about 4–8 meters tall, and both the leaves and berries are harvested for their nutritional and medicinal properties. Like many other species within the Myrtaceae family, camu camu evolved within the Amazon Basin, and grows along the rocky riverbeds and swamps in Brazil, Venezuela, Peru, Colombia, and Bolivia. There are significant gaps in knowledge in the phylogenetic history of the plant, which creates avenues for future research. In terms of phenology, camu camu plants synchronize their growth cycles according to the flood-levels of these watersheds, producing small bisexual white flowers during periods with low water levels. As the water levels rise, the plant transitions into its fruiting stage, where it produces small, reddish-purple fleshy fruits. The camu camu berries are known for their high vitamin C content, and small amounts of various amino acids and fatty acids. The berries of camu camu have many anti-inflammatory and antimicrobial properties, and they are a valuable food source to the birds, mammals, fish, and people living in the Amazon Basin. The growing scientific studies on the plant's medicinal properties have increased demand for the fruit globally. The growth in international markets has led farmers to harvest more camu camu berries, negatively impacting wild populations. By Nicole Phelan and Holland Smith #CamuCamu
23. Wolffia Globosa: The Super Small Superfood Wolffia globosa, which is commonly known as mankai, Asian watermeal, and duckweed, is a species of flowering plant that is currently considered to be the smallest known flowering plant! Wolffia globosa is a genus from the lemnaceae family native to Asia as well as pacific coasting countries. Wolffia grows in groups on the surface of lakes and large bodies of calm water, ponds, and even marshes together with other aquatic plants and life. Due to its rapid growth rate and the relative ease at which it grows on calm bodies of water, Wolffia globosa was traditionally only harvested, and not cultivated by indigenous peoples. Wolffia was initially discovered by William Roxburg, a Scottish botanist and surgeon practicing in parts of India in the late 1780s to early 1800s, in 1832. In 1984, Wayne P. Armstrong, nicknamed “Mr. Wolffia”, became the first individual to discover Wolffia Globosa in California while he was studying duckweeds from a lake nearby his lab near the San Dieguito river (Armstrong, 2021). Wolffia globosa is an extremely versatile plant. Initially, it was used by indigenous peoples as a protein packed vegetable, and it is still being used in traditional Thai cuisine. Due to the extremely fast growth rate of Wolffia globosa, indigenous peoples were also able to repurpose mankai, using it as feed for their livestock. Currently, Wolffia globosa has found itself at the center of superfood trends due to its phenomenal nutritional benefits. Outside of the culinary world, Wolffia globosa can be found at the center of research regarding new biofuels and potential bioremediation strategies to combat pollution. By Akshay Chellappa, Brian Estarella-Murphy, #Mankai
24. Ulluco: Resilience and Preservation Through Time Ullucus tuberosus (ulluco) is a crop grown throughout South America for its starchy tubers and spinach-like leaves. Grown alongside humans for thousands of years, the genetic diversity of ulluco along with indigenous knowledge of the crop has enabled it to survive amidst challenges of Spanish colonial rule. One challenge was competition from introduced European cultivars such as barley and carrots. Another challenge was that Indigenous groups who cultivativated ulluco faced hardships and challenges throughout colonization that echo into modern times. Despite these barriers, Indigenous Andean groups continued cultivating ulluco on smaller scales. This preserved Indigenous agricultural methods, and therefore ulluco biodiversity, still today. Now, ulluco has been proposed as a crop that could help solve diminishing food supplies amidst climate change. By Audrey Hernandez, Justin Haggard #Ulluco
25. From medicine to industry: the multifaceted history of Guaraná (Paullinia cupana Kunth., Sapindaceae) Paullinia cupana, nature’s caffeine tablet, is a climbing shrub with a very deep history within indigenous South American tribes. Known by locals as guaraná, it has an easily recognizable fruit with an eyeball-like structure, giving it the name of “eye of the gods.” The seeds contain the highest concentrations of caffeine of any other plant – about four times the amount found in coffee beans. It was commonly used by the Sateré Maué people to make beverages and used as medicine for bodily discomfort as far back as over 2,000 years ago. Nowadays, despite its popularity, the only country that cultivates guaraná is Brazil, with over 15,000 hectares of land growing it. Its importance to Brazilians has led to the deforestation of parts of the Amazon in order to make room to grow guaraná. In 2006, the cash value of domestic production of guaraná was over $13 billion and has continued to increase due to the growing domestic and international demand. Due to the seed’s high caffeine content and unique flavor, its extract has become a widely used additive in energy drinks. The most recognizable of which is the very popular soda, Guaraná Antarctica, and is the biggest competitor of Coca Cola in Brazil. By Tim Sisneros and Edward Teng.
26. Lorem Ipsum: The Fake Latin Text Substitute Elementum facilisis leo vel fringilla est ullamcorper eget nulla facilisi etiam dignissim diam quis enim lobortis scelerisque fermentum dui faucibus in ornare quam viverra orci sagittis eu volutpat odio facilisis mauris sit amet massa vitae tortor condimentum lacinia quis vel eros donec ac odio tempor orci dapibus ultrices in iaculis nunc sed augue lacus viverra vitae congue eu consequat ac felis donec et odio pellentesque diam volutpat commodo sed egestas egestas fringilla phasellus faucibus scelerisque eleifend donec pretium vulputate sapien nec sagittis aliquam malesuada bibendum arcu vitae elementum curabitur vitae nunc sed velit dignissim sodales ut eu sem integer vitae justo eget magna fermentum iaculis eu non diam phasellus vestibulum lorem sed risus ultricies tristique nulla aliquet enim tortor at auctor urna nunc id cursus metus aliquam eleifend mi in nulla posuere sollicitudin aliquam ultrices sagittis orci a scelerisque purus semper eget duis at tellus at urna condimentum mattis pellentesque id nibh tortor id aliquet lectus proin nibh nisl condimentum id venenatis a condimentum vitae sapien pellentesque habitant morbi tristique senectus et netus et malesuada fames ac turpis egestas sed tempus urna et pharetra pharetra massa massa ultricies mi quis hendrerit dolor magna eget est lorem ipsum dolor sit amet consectetur adipiscing elit pellentesque habitant morbi tristique senectus et netus et malesuada fames ac turpis egestas integer eget aliquet nibh praesent tristique magna sit amet purus gravida quis blandit turpis cursus in hac habitasse platea dictumst quisque sagittis purus sit amet volutpat consequat mauris nunc congue nisi vitae suscipit tellus. By Lorem Ipsum #lorem #ipsum

Aronia melanocarpa: fine wines to phenols

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